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R.  Rosengren, Chautems, D., Cherix, D., Fortelius, W., and Keller, L., ? Separation of two sibling species of Formica L. ants by a behavioural choice test based on brood discrimination.?, Mem. Zool., vol. 48, pp. 237-249, 1994.

R.  Rosengren, Fortelius, W., Lindström, K., and Luther, A., ?Phenology and causation of nest heating and thermoregulation in red wood ants of the Formica rufa group studied in coniferous forest habitats in southern Finland.?, Annales Zoologici Fennici, vol. 24, pp. 147-155, 1987.

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R.  Rosengren and Fortelius, W., ? Trail communication and directional recruitment to food in red wood ants (Formica). ?, Annales Zoologici Fennici, vol. 24, pp.  137-146, 1987.

R.  Rosengren and Fortelius, W., ?Ortstreu in foraging ants of the Formica rufa group ? hierarchy of orienting cues and long-term memory. ?, Insectes Sociaux, vol. 33, pp. 306-337, 1986.

L.  Passera, Keller, L., Grimal, A., Chautems, D., Cherix, D., Fletcher, D. J. H., Fortelius, W., Rosengren, R., and Vargo, E. L., ?Carbohydrates as a energy source during the flight of sexuals of the ant Formica lugubris (Hymenoptera; Formicidae)?, Entomol. Gener., vol. 15, pp. 25-32, 1990.

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W.  Fortelius, Rosengren, R., Cherix, D., and Chautems, D., ?Queen recruitment in a highly polygynous supercolony of Formica lugubris (Hymenoptera: Formicidae).?, Oikos, vol. 57, pp. 193-200, 1993.

W.  Fortelius, Pamilo, P., Rosengren, R., and Sundström, L., ?Male size dimorphism and alternative reproductive tactics in Formica exsecta ants (Hymenoptera;Formicidae).?, Ann. Zool. Fennici, vol. 24, pp. 45-54, 1987.

W.  Fortelius, ?Mating behavior in the polygynous/polydomous wood ant Formica aquilonia.?, Ann. Zool. Fennici , vol. 42, pp. 213-224., 2005.

W.  Fortelius, ?Mating and phototaxis of wood ant, Formica lugubris Zett. (Hymenoptera, Formicidae).?, Memorabilia Zoologica, vol. 48, pp. 69-80, 1994.

D.  Cherix, Fletcher, D. J. C., Chautems, D., Fortelius, W., Gris, G., Keller, L., Rosengren, R., Vargo, E. L., and Walther, F., ?Attraction of sexes in Formica lugubris Zett. (Hymenoptera, Formicidae).?, Insectes Sociaux, vol. 40, pp. 319-32, 1993.

D.  Cherix, Chautems, D., Fletcher, D. J. C., Fortelius, W., Gris, G., Keller, L., Passera, L., Rosengren, R., Vargo, E. L., and Walther, F., ?Alternative reproductive strategies in Formica lugubris Zett. (Hymenoptera, Formicidae). ?, Ethol. Ecol. Evol., Special Issue , vol. 1, pp. 61-66, 1991.